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The success of arthropods derives in large part from the evolution of their unique, nonliving, organic, jointed exoskeleton (see figure), which not only functions in support but also provides protection and, with the muscle system, contributes to efficient locomotion. The exoskeleton is composed of a thin, outer protein layer, the epicuticle, and a thick, inner, chitin–protein layer, the procuticle. In most terrestrial arthropods, such as insects and spiders, the epicuticle contains waxes that aid in reducing evaporative water loss. The procuticle consists of an outer exocuticle and an inner endocuticle. In the exocuticle there is cross-bonding of the chitin–protein chains ...(100 of 5803 words)