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In the following century the work of Aristotle, regarded as the first great biologist, was of inestimable value to medicine. A pupil of Plato at Athens and tutor to Alexander the Great, Aristotle studied the entire world of living things. He laid what can be identified as the foundations of comparative anatomy and embryology, and his views influenced scientific thinking for the next 2,000 years. After the time of Aristotle, the centre of Greek culture shifted to Alexandria, where a famous medical school was established about 300 bce. There the two best medical teachers were Herophilus, whose treatise on anatomy ...(100 of 21288 words)